Every year, the KND choose someone to be "IT" which involves a game of tag to choose the next Supreme Leader, so Numbuh 362 sets up the game after she decides to resign after being tired of the job. But when Numbuh 2 accidentally tags Father, he becomes Supreme Leader.
Now, it's up to Numbuh 1 and Numbuh 362 to stop him before he wipes out the entire KND.
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